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TheDrosophila boulegene is expressed exclusively in the male germline and encodes an RNA binding protein closely related to the mammalian fertility factors encoded by theDAZ(Deleted in Azoospermia) andDAZL(DAZ-like) genes. Mutation ofbouleblocks both meiotic divisions. Differentiation nonetheless continues, resulting in tetraploid spermatids that fail to mature into sperm. We have found that Boule...