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The biosynthesis pathways of coenzyme A (CoA) in most archaea involve several unique enzymes including dephospho‐CoA kinase (DPCK) that converts dephospho‐CoA to CoA in the final step of CoA biosynthesis in all domains of life. The archaeal DPCK is unrelated to the analogous bacterial and eukaryotic enzymes and shows no significant sequence similarity to any proteins with known structures. Unusually,...