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SECIS elements recode UGA codons from “stop” to “sense.” These RNA secondary structures, present in eukaryotic selenoprotein mRNA 3′ untranslated regions, recruit a SECIS binding protein, which recruits a selenocysteine-specific elongation factor-tRNA complex. Elucidation of the assembly of this multicomponent complex is crucial to understanding the mechanism of selenocysteine incorporation. Coprecipitation...