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In Saccharomyces cerevisiae, repair of insertion/deletion loops is carried out by Msh2–Msh3-mediated mismatch repair (MMR). Msh2–Msh3 is also required for 3′ non-homologous tail removal (3′ NHTR) in double-strand break repair. In both pathways, Msh2–Msh3 binds double-strand/single-strand junctions and initiates repair in an ATP-dependent manner. However, the kinetics of the two processes appear different;...