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Selective recognition and transport of Na+ and Ca2+ ions by sodium–calcium exchanger (NCX) proteins is a primary prerequisite for Ca2+ signaling and homeostasis. Twelve ion‐coordinating residues are highly conserved among NCXs, and distinct NCX orthologs contain two or three carboxylates, while sharing a common ion‐exchange stoichiometry (3Na+:1Ca2+). How these structural differences affect the ion‐binding...
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