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Redox, optical and kinetic characteristics of the four-heme cytochrome c tightly bound to the reaction center complexes of Rhodopseudomonas viridis have been studied. The two high-potential hemes, previously thought to be identical, are shown to differ in midpoint potentials, absorption spectra and kinetics of photooxidation. One heme is characterized by E m = 380 +/- 10 mV, and a split α-band...