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The heterotrimeric UmuD' 2 C complex of Escherichia coli has recently been shown to possess intrinsic DNA polymerase activity (DNA pol V) that facilitates error-prone translesion DNA synthesis (SOS mutagenesis). When overexpressed in vivo, UmuD' 2 C also inhibits homologous recombination. In both activities, UmuD' 2 C interacts with RecA nucleoprotein filaments. To examine...