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In 2005, soybean plants with severe mosaic and leaf rugose symptoms were found in Japan. The plants contained flexuous and filamentous virus-like particles with a modal length of 500 nm. Double-stranded RNA analysis revealed a single major band of ~7000 bp. Reverse transcription-PCR for Potyviridae diagnosis generated a 1.5-kbp cDNA fragment, which contained a partial NIb gene, a coat protein (CP)...