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The hyperpolarization‐activated cyclic‐nucleotide‐gated (HCN) ion channels control nerve impulse transmission and cardiac pacemaker activity. The modulation by cAMP is critical for the regulatory function of HCN in both neurons and cardiomyocytes, but the underlying mechanism is not fully understood. Here, we show how the structure of the apo cAMP‐binding domain of the HCN4 isoform has contributed...
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