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Axonemal dynein is an ATP‐dependent microtubular motor protein responsible for cilia and flagella beating, and its dysfunction can cause diseases such as primary ciliary dyskinesia and sperm dysmotility. Despite its biological importance, structure‐based mechanisms underlying axonemal dynein motors remain unclear. Here, we determined the X‐ray crystal structure of the human inner‐arm dynein‐d (DNAH1)...