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Metriorhynchoid thalattosuchians were a marine clade of Mesozoic crocodylomorphs that evolved from semi‐aquatic, “gharial”‐like species into the obligately pelagic subclade Metriorhynchidae. To explore whether the sensory and physiological demands of underwater life necessitates a shift in rostral anatomy, both in neurology and vasculature, we investigate the trigeminal innervation and potential somatosensory...