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Until about 25 years ago, classification of the flatfishes was based on the Regan-Norman model. Beginning with the work of Amaoka in 1969, the order started to be analysed in finer detail, and in the 1980s many questions arose concerning flatfish phylogeny as expressed in the Regan-Norman model. Two of the major questions were whether the order is monophyletic and which groups within the order are...