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Defective influenza A virus RNAs analyzed in two studies so far possess at least 80–90 nucleotides from the 5′ end of the virion RNA segment and more typically around 200 nucleotides, whereas the 3′ sequence could be as short as 25 nucleotides (P. A. Jennings et al., Cell 34, 619–627; 1983; S. D. Duhaut and N. J. Dimmock, Virology 247, 241–253, 1998). To determine the biological significance of the...