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Voltage-dependent anion channels (VDACs) are the porins in the outer mitochondrial membrane allowing metabolite flux between mitochondria and the cytoplasm. The permeabilities of the VDACs to ATP−4, ADP−3, creatine phosphate2−, Pi2−, Pi−, and other charged metabolites depend on the membrane potential. But neither the existence of the electrical potential across the outer membrane of mitochondria,...
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