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Tinbergen is famous for emphasizing behavioral fieldwork and experimentation under natural circumstances, for founding the field of ethology, for getting a Nobel Prize, and for mentoring Richard Dawkins. He is known for dividing behavior studies into physiology, development, natural selection, and evolutionary history. In the decades since Tinbergen was active, some of the best research in animal...
Niko Tinbergen's (Zeit. Tier. 20, 1963, 410) paper ‘On aims and methods of ethology’ is appropriately remembered as the paper in which Tinbergen characterized ethology as ‘the biological study of behavior’ and went on to explain that to study behavior biologically is to ask four distinct questions about it: (1) How is it caused physiologically? (2) What is its survival value? (3) How has it evolved?...
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