During the last several years Professor Rozenberg and I have been working closely together to try to unravel the secrets of complex DNA manipulations in ciliates. His creative and incisive analysis has given us a fresh perspective on the remarkable acrobatics that ciliates perform with their DNA gene sequences. I am pleased to contribute this paper on the structure of germline genes, which ciliates process in phenomenal ways, as an expression of my appreciation and admiration for his intelligence and energy in the quest to explain the extraordinary DNA gymnastics that the tiny ciliates perform.