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Electrostatic considerations predict that the binding of divalent cations to singly charged acidic lipids is likely to proceed with a 1:2 ion-to-lipid stoichiometry, but the fact that divalent cations at ∼0.1 M concentrations reverse the sign of the negative surface potential of membranes provides evidence for 1:1 stoichiometry. In this work, analytic expressions are obtained for 1:1 and 1:2 binding...
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