A pedal phalanx of Homo from the Paleolithic stratum 11 at Denisova Cave, Altai, is described. A manual phalanx, from which DNA of a hominin belonging to a hitherto unknown species was extracted, had been found in the same layer. The new specimen has been identifi ed as a left proximal phalanx of the fourth or fi fth pedal digit, apparently that of an adult. The bone is rather elongated and has a very robust and broad diaphysis. While being generally archaic in appearance, the specimen demonstrates certain traits that are intermediate between the Neanderthal and the early modern morphology. The dorso-proximal orientation of the metatarsal facet of the base indicates a peculiar gait, resembling race walking.