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Puccinia metanarthecii and P. pachycephala differed in their heteroecious life cycle. The former rust fungus formed spermogonial/aecial stages on Ixeris dentata and uredinial/telial stages on Metanarthecium luteo-viride or Aletris foliata. The latter formed spermogonial/aecial stages on Ligularia hodgsonii and uredinial/telial stages on Veratrum maackii var. longibracteatum and V. maackii var. parviflorum...
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