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The proton reactivity of a soil bacterium, Pseudomonas putida ATCC12633 was investigated in two physiologically different states: (i) as free-living cells, and (ii) as a 5-day biofilm formed in a sandy column.Acid–base data analysis and modeling based on a three-site non-electrostatic model showed that biofilm has a proton exchange capacity 3.8 times higher than that of free-living cells (13.2mmol/g...
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