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T cells recognize antigens at the two‐dimensional (2D) interface with antigen‐presenting cells (APCs), which trigger T‐cell effector functions. T‐cell functional outcomes correlate with 2D kinetics of membrane‐embedded T‐cell receptors (TCRs) binding to surface‐tethered peptide‐major histocompatibility complex molecules (pMHCs). However, most studies have measured TCR–pMHC kinetics for recombinant...
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