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Elongation factor G (EF‐G) plays a crucial role in two stages of mRNA‐(tRNA)2 translocation. First, EF‐G•GTP enters the pre‐translocational ribosome in its intersubunit‐rotated state, with tRNAs in their hybrid (P/E and A/P) positions. Second, a conformational change in EF‐G's Domain IV induced by GTP hydrolysis disengages the mRNA‐anticodon stem‐loops of the tRNAs from the decoding center to advance...
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