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Transcription initiation by bacterial σ 54 -RNA polymerase requires a conformational change of the holopolymerase–DNA complex, driven by an enhancer-binding protein. Although structures of the core polymerase and the more common σ 70 factor have been determined, little is known about the structure of the σ 54 variant. We report here the structure of an Aquifex aeolicus σ ...
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