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His354 and His358, two highly conserved histidines in Xenopus laevis (6-4) photolyase [equivalent to His401 and His405, in Dunaliella salina (6-4) photolyase], are critical for photoreactivation. They act as a base and an acid, respectively. However, the remaining high repair activity when the pH value is higher than the pKa of histidine suggests the involvement of other basic amino acids in photoreactivation...
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