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The first high-resolution structure of a P-type ATPase, that of the Ca 2+ -ATPase of skeletal muscle sarcoplasmic reticulum, was published in 2000. This structure has provided many clues to how the Ca 2+ -ATPase might work, but no complete answers. The Ca 2+ -ATPase structure reveals no clear pathway from the cytoplasmic side of the membrane to the pair of high-affinity...
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