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The aa 3 -type cytochrome c oxidases (CytcOs) from e.g. Rhodobacter sphaeroides and Paracoccus denitrificans harbor two proton-transfer pathways. The K pathway is used for proton uptake upon reduction of the CytcO, while the D pathway is used after binding of O 2 to the catalytic site. The aim of the present study was to determine whether or not CytcO in which the K pathway is blocked...
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