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The Aktulagay section in Kazakhstan provides an expanded northern Tethyan record of the middle Ypresian (calcareous nannoplankton zones NP11–13, ~54–50Ma), including the Early Eocene Climatic Optimum (EECO). The marl sequence features a series of sapropel beds, observed throughout the Peri-Tethys, indicative of the basin-wide occurrence of episodic hypoxic events. In order to unravel the paleoenvironmental...
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