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In Rhodobacter sphaeroides, transfer of the first electron in quinol oxidation by the bc 1 complex shows kinetic features (a slow rate (approx. 1.5x10 3 /s), high activation energy (approx. 65 kJ/mol) and reorganization energy, λ (2.5 V)) that are unexpected from Marcus theory and the distances shown by the structures. Reduction of the oxidized iron-sulfur protein occurs after formation...
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