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In contrast to seed plants, the gametophytes of seed‐free plants develop pluripotent meristems, which promote and sustain their independent growth and development. To date, the cellular basis of meristem development in gametophytes of seed‐free ferns remains largely unknown. In this study, we used Woodsia obtusa, the blunt‐lobe cliff fern, to quantitatively determine cell growth dynamics in two different...