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The present study consists of the microscopic and geochemical analyses of specimens of the trace fossil Phymatoderma from the Pliocene deep-marine strata (Shiramazu Formation, Japan). The aim of this paper is to assess whether benthic-pelagic coupling functioned in the ancient ocean as it does in the modern ocean. Various types of microfossils (e.g., coccoliths, diatoms, planktonic foraminifera, and...
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