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The Caenorhabditis elegans vulva has served as a paradigm for how conserved developmental pathways, such as EGF–Ras–MAPK, Notch and Wnt signaling, participate in networks driving animal organogenesis. Here, we discuss an emerging direction in the field, which places vulva research in a quantitative and microevolutionary framework. The final vulval cell fate pattern is known to be robust to change,...
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