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Spinach thylakoids were resuspended in a buffer including either 200 mM sucrose or 100 mM KCl. singleturnover flash excitation and the luciferin-luciferase system were used to energize and follow ATP formation. The effect of the permeable buffer pyridine was measured on the ATP formation onset lag and on the post-illumination ATP yield. Consistent with a bulk phase delocalized proton gradient coupling...
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