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In angiosperms, pollen tube growth is critical for double fertilization and seed formation. Many of the factors involved in pollen tube tip growth are unknown. Here, we report the roles of pollen‐specific GLYCEROPHOSPHODIESTER PHOSPHODIESTERASE‐LIKE (GDPD‐LIKE) genes in pollen tube tip growth. Arabidopsis thaliana GDPD‐LIKE6 (AtGDPDL6) and AtGDPDL7 were specifically expressed in mature pollen grains...