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Quercetin 2,4‐dioxygenase (quercetinase) from Streptomyces uses nickel as the active‐site cofactor to catalyze oxidative cleavage of the flavonol quercetin. How this unusual active‐site metal supports catalysis and O2 activation is under debate. We present crystal structures of Ni‐quercetinase in three different states, thus providing direct insight into how quercetin and O2 are activated at the Ni...
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