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Phage λ DNA packaging is accompanied by prohead expansion, due to structural changes in gpE, the major capsid protein. Rearrangement of the gpE lattice creates binding sites for trimers of gpD, the head stabilization protein. λ-Like phage 21's shp gene is homologous to λ's D gene. gpD and gpShp share 49% amino acid identity. To ask whether gpShp could stabilize the λ head shell, we replaced λ's D...
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