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Protochlorophyllide oxidoreductase (POR), which converts protochlorophyllide into chlorophyllide, is the only light‐dependent enzyme in chlorophyll biosynthesis. While its catalytic reaction and importance for chloroplast development are well understood, little is known about the post‐translational control of PORs. Here, we show that cpSRP43 and cpSRP54, two components of the chloroplast signal recognition...