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Early spermatophytes have been discovered at Red Hill, a Late Devonian (Famennian) fossil locality in north-central Pennsylvania, USA. The Red Hill locality contains an Archaeopteris-dominated flora within an outcrop of the Duncannon Member of the Catskill Formation. Palynological analyses of the plant fossil-bearing horizons within the Red Hill outcrop indicate deposition within the VCo palynozone...
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