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The basic helix-loop-helix transcription factor Twist1 is essential for normal limb development. Twist1 −/− embryos die at midgestation. However, studies on early limb buds found that Twist1 −/− mutant limb mesenchyme has an impaired response to FGF signaling from the apical ectodermal ridge, which disrupts the feedback loop between the mesenchyme and AER, and reduces and shifts anteriorly...
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