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Primases synthesize short RNA strands on single-stranded DNA templates, thereby generating the hybrid duplexes required for the initiation of synthesis by DNA polymerases. We present the crystal structure of the catalytic unit of a primase enzyme, that of a ∼320 residue fragment of Escherichia coli primase, determined at 2.9 Å resolution. Central to the catalytic unit is a TOPRIM domain that is strikingly...
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