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Deep-sea trace fossils of the Nereites ichnofacies occur in the uppermost Kimmeridgian–Berriasian deep-water turbidite deposits of the Kostel Formation in SW Bulgaria. The trace fossil assemblage contains Tithonian Ophiomorpha isp. and Ophiomorpha annulata, which are the oldest deep-sea representatives of this ichnogenus. They indicate much older invasion of Ophiomorpha into deep-sea, which was hitherto...
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