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Savannah (C 4 ) grasses are first recorded at low latitudes in the mid-Miocene prior to their expansion towards mid-latitudes by approximately the Miocene–Pliocene boundary. In an attempt to determine the timing of the spread of savannah grasses into the South African highveld, a palaeoecological study of some of the oldest faunal deposits in the region (mid-Pliocene) was undertaken. The combination...
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