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The endomembrane system consists of various membrane‐bound organelles including the endoplasmic reticulum (ER), Golgi apparatus, trans‐Golgi network (TGN), endosomes, and the lysosome/vacuole. Membrane trafficking between distinct compartments is mainly achieved by vesicular transport. As the endomembrane compartments and the machineries regulating the membrane trafficking are largely conserved across...