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The interaction between CD4 and major histocompatibility complex (MHC) class II proteins is critical for the activation of CD4 + T cells, which are involved in transplantation reactions and a number of autoimmune diseases. It is known that the CD4 N-terminal immunoglobulin variable region-like domain (D1) is directed toward and reaching into the two membrane-proximal domains of the MHC class...
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