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When monitored by 1 H NMR at various pH values, most of the C-2 proton signals from 12 His residues of the isolated β subunit of thermophilic F 1 -ATPase (TF 1 ) could be separately observed. Two of them were assigned to His-179 and His-200 which reside at the entrance of a conical tunnel to reach catalytic site in the crystal structure of F 1 -ATPase. His-200...
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