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In 1957, a unique pattern of hydrogen bonding between N3 and O4 on uracil and N7 and N6 on adenine was proposed to explain how poly(rU) strands can associate with poly(rA)‐poly(rU) duplexes to form triplexes. Two years later, Karst Hoogsteen visualized such a noncanonical A–T base‐pair through X‐ray analysis of co‐crystals containing 9‐methyladenine and 1‐methylthymine. Subsequent X‐ray analyses of...
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